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[ Amanita franchetii f. lactella (E.-J. Gilbert & Kühner) Bon & Contu in Contu ]

Amanita franchetii (Boud.) Fayod
" Franchet's Amanita"


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Technical description (t.b.d.)

BRIEF DESCRIPTION: The following is largely based on the description by Neville and Poumarat (2004).

The cap of Amanita franchetii is 65 - 80 mm wide, citron yellow to yellow, often palest towards the margin, sometimes whitish, hemispheric then convex, finally planar, often rather irregular, sometimes depressed in the center in age, shiny, viscid then silky, not virgate, with an appendiculate margin, sometimes striate in age.  The volva is present as small, generally pyramidal warts, sulfur-yellow becoming paler and sordid (for example, pale grayish ochre), easily removable, densely placed.  The flesh is 5 mm thick above the stem, white, with a thin yellow region just below the cap skin, tending to turn brown when wounded.

The gills are free at maturity, distant, white, up to 7 mm broad, not or only finely floccose.  The short gills are truncate, rounded truncate, or attenuate.

>The stem is 65 - 80 × 18 - 20 mm, white, solid, firm, striate above the ring, smooth or slightly flocculose below the ring, with a napiform bulb up to 33 mm wide.  The ring is membranous, white, striate on the upper surface, with yellow volval warts on the edge.  The volva is present as 2 or 3 incomplete rings of warts at the top of the bulb, at first yellow becoming yellowish to pale grayish-beige with age.  The flesh is white, tending to turn brown when wounded especially at the base of the stem and the bulb.

The spores measure 7.5 - 9.5 (-11) × (5-) 5.5 - 7 (-7.5) µm and are broadly ellipsoid to ellipsoid and amyloid.  Clamps are absent at bases of basidia.

This species was originally described from France but has been reported from much of Europe and northern Africa.  This species occurs in the same environments with its form queletii.  It is associated with woody plants: oaks (Quercus pubescens, Q. suber), chestnut (Castanea sativa), and pine (Pinus).

Neville and Poumarat accept two forms of the present species in addition to the type form: f. lactella and f. queletii.

The name "A. franchetii" is mistakenly applied to an otherwise undescribed taxon that is commonly found in California.  The name "Amanita apera" [for example, see H. D. Thiers (1982)] has also been applied to both the European and North American taxa; however, the latter epithet applies to a lepiotaceous fungus.  -- R. E. Tulloss and L. Possiel

Amanita franchetii f. lactella (E.-J. Gilbert & Kühner) Bon & Contu in Contu.

[Note: Differing from f. franchetii by having a milk white cap rather than one which ranges from citron yellow to whitish. In the most recent restatement of information concerning f. lactella, Neville and Poumarat do not make a case for segregating taxonomically pure white specimens from specimens that are almost white. It is worth noting that we have found no value in segregation of palely pigmented and albino specimens of a number of taxa (for example, A. brunnescens G. F. Atk., A. caesarea (Scop. : Fr.) Pers., and A. phalloides (Fr. : Fr.) Link, etc.).  In other cases, RET has found some pallid and white infraspecific taxa to be distinct at the ranks of variety or species (for example, A. rubescens var. alba Coker).  A careful evaluation of the status of f. lactella would be valuable -- especially since a neotype has been designated.]

The following is based on the description of Bertault based on north African material associated with Quercus suber which is provided in full by Neville and Poumarat (2004).

The cap of Amanita franchetii f. lactella is 70 - 90 mm wide, pure white, becoming a little brownish in the center in age, globose or hemispheric at first, then convex, sometimes with a little umbo, finally planar or subdepressed, matte, subviscid when moist, with a smooth, nonappendiculate and nonstriate margin, except in age.  The volva is present as membranous, densely arranged, pyramidal warts in the center of the cap, bright golden yellow, easily removable, cleanly detached from browning areas; the warts are sparser, flatter, and paler towards the edge.  The flesh is white.

The gills are crowded, free to distant from the stem, white, narrowing in both directions, up to 8 mm broad, with a finely floccose edge.  The short gills are truncate.

The stem is 90 - 100 × 10 - 20 mm, solid, subcylindric, flaring at the top, white, slightly squamulose below the ring, stained brownish at the bottom of the stem, with a slightly spindle-shaped bulb.  The ring is membranous, placed high on the stem, skirt-like, finely striate above, with an irregularly splitting and rather thickened edge, and tinted bright yellow or light brown on the edge.  The volva is present as vague circles of yellow or in mature material as an irregular a bistre line around the top of the bulb, but the latter is not always present.  The flesh is white, turning brownish in the bulb when cut.

The odor is indistinct and the taste is sweet.

The spores measure 7.5 - 8.5 (-9) × 4.8 - 5 µm and are ellipsoid to elongate and amyloid.  Clamps are absent at bases of basidia.  Bertault's measurements describe spores proportionately narrower than those of f. franchetii.  In contrast, Neville and Poumarat cite measurements on 20 spores from French material which they considered otherwise in poor condition, and those measurements are 7.5 - 9 (-9.5) × 6 - 7 and are broadly ellipsoid to ellipsoid and amyloid.

This taxon occurs in Spain, France, Italy, and Morocco in association with oak (Quercus suber, Q. robur) and Carpinus betulus. -- R. E. Tulloss and L. Possiel

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Last changed 26 April 2009.
This page is maintained by R. E. Tulloss.
Copyright 2006, 2007, 2008, 2009 by Rodham E. Tulloss.